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蓝紫白鬼伞

Leucocoprinus ianthinus

 

(Sacc.) P. Mohr 1994

基本信息 General Info.

  • 通用中文名Chinese name:蓝紫白鬼伞
  • 拉丁学名Scientific name:Leucocoprinus ianthinus (Sacc.) P. Mohr 1994
  • 科Family:蘑菇科Agaricaceae
  • 属Genus:白鬼伞属Leucocoprinus
  • 分布Distribution:广泛分布于各地。
  • 有分布的省区Provinces:*暂无信息*
  • 生态Ecology:出现于花盆、温室等人工环境中。
  • 营养类型Nutrition Type:腐生Saprotrophic: 分解死亡生物体或有机物获取养分
  • 置信指标Confidence:★★✩

物种介绍

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来源:心不狠站不稳

蓝紫白鬼伞是一种常见于花盆、温室等人工环境的蘑菇。它并非我国本土菌种,而很可能潜伏在土中,随营养土和观赏植物交易传入我国。其原产地可能在热带地区。

主要特征

担子果小型,常多个生长;菌盖幼时近卵形,后开展为钟形、斗笠形或近平展,中部略凸起,边缘有长辐射状棱纹;盖表密被大量褐色至紫褐色细鳞片,鳞片在中部形成一整块,朝四周渐稀;菌褶白色,老后可能变色,离生;菌柄近白色,中上部有菌环,基部有紫褐色鳞片;褶缘囊状体幼年个体明显,老化后不明显(Gierczyk & Dubiel 2014);菌盖鳞片由排列成不规则链状的球形或椭圆形细胞组成;担孢子芽孔有盖。(Stallman 2019, Vellinga 2001, Gierczyk & Dubiel 2014)

分类学争议

Leucocoprinus lilacinogranulosus是一个相似的物种。主流学界一般认为两者是同一物种,但有观点认为两者不同。

Locquin (1945)认为L. lilacinogranulosus显微镜下菌盖表皮膜皮型,而L. ianthinus菌盖表皮呈“丝状(filamentous)”。Reid (1989)后续对L. ianthinus模式标本的研究,证明Locquin的“丝状”描述是不准确的,两个物种的微观特征其实相当类似;但他指出两者宏观上存在差异——L. ianthinus菌盖无明显沟纹,展开后中部凸起,整个菌盖被辐射状的纤维状鳞片;L. lilacinogranulosus则有明显沟纹,展开后无明显凸起,菌盖中部被雀斑状的细鳞片。Reid既未支持也未反对合并两物种。Bon (1993)后续合并了两物种,但他怀疑孢子大小和菌核的存在与否(L. lilacinogranulosus观察到菌核)可能暗示不同物种。多数学者,如Vellinga (2011)和主流数据库现在支持合并两物种。Gierczyk & Dubiel 2014则支持两者不同,理由包括L. lilacinogranulosus边缘有沟纹、紫色鳞片颗粒状、菌柄基部带紫色调、孢子(7)8–10.5(11) × (5.5)6–7.5(8) μm;而L. ianthinus边缘平滑、紫色鳞片纤维状、菌柄基部无紫色调、孢子6.5–10 × 5.75–6.5 μm。

相似物种

L. heinemannii为代表的一些物种也有紫色调的鳞片,看上去和蓝紫白鬼伞相似(这是一个形态学上十分相似但系统树上较分散的复合群,尤其是L. parvipileus这个物种尤和本种相似)。这些物种和本种的区别是:这些物种的孢子上没有芽孔,而真正的蓝紫白鬼伞的孢子上有一个有盖芽孔(Stallman 2019, Vellinga 2001, Gierczyk & Dubiel 2014)。另外,复合群内物种的孢子长度普遍小于7.5μm,宽度普遍短于4.5μm,而蓝紫白鬼伞的孢子长度虽然有多种记载,但基本都超过这些数据,例如Arun Kumar & Manimohan 2009记载8–11 × 5.5–7μm、Gierczyk & Dubiel 2014记载8.0-11.0 × 6.2-7.5 μm、Ludwig 2012记载(7)8-10.5(11) × (5.5)6-7.5(8)μm。关于我国的个体,作者对家中长出来的样本的孢子进行了粗略测量,数据大约是7-9×5-7.5μm。目前不排除我国的“蓝紫白鬼伞”中混有此复合群的物种。

褐顶白鬼伞L. brebissonii和本种也有些相似,但可以明确区分。主要区别是褐顶白鬼伞(除非老化后)鳞片通常更偏黑褐色(颜色区别在幼年担子果上尤为明显),据中国真菌志记载芽孔一般缺乏孔盖,且它通常不会出现在花盆中。Gierczyk & Dubiel 2014记载褐鳞白鬼伞的孢子“have a distinct papilla-like apex”(有一个乳突状的顶部),但中国真菌志的记载中未见。Arun Kumar & Manimohan 2009的检索表中记载L. ianthinus的菌柄受伤变色(而褐顶白鬼伞不会),但在我采集的样本上未见此特点(至少不明显)。

物种介绍-参考文献
1. Leucocoprinus lilacinogranulosus (Henn.) Locq. in Poland. 2014. Błażej Gierczyk & Grzegorz Dubiel. Acta Mycologica 49(1):59-67    https://doi.org/10.5586/am.2014.005
2. Three new cryptic Caribbean species in the Leucocoprinus heinemannii complex (Agaricaceae, Agaricales). 2020. Alfredo Justo & Claudio Angelini & Alberto Bizzi & Alessia Tatti & Alfredo Vizzini. Mycological Progress, 19(12), 1445–1457    https://doi.org/10.1007/s11557-020-01638-9
3. The genera Leucoagaricus and Leucocoprinus (Agaricales, Basidiomycota) in Kerala State, India. 2009. T.K. Arun Kumar, Patinjareveettil Manimohan. Mycotaxon 108(1):385-428    https://doi.org/10.5248/108.385
4. 中国真菌志 第52卷 环柄菇卷(蘑菇科). 2019. 杨祝良主编. 北京:科学出版社    
5. Taxonomy of Hawaiʻi Island’s Lepiotaceous (Agaricaceae) Fungi: Chlorophyllum, Cystolepiota, Lepiota, Leucoagaricus, Leucocoprinus. 2019. Jeff Stallman. Thesis for: Master of ScienceAdvisor: Michael Shintaku, Don Hemmes, Nicole Hynson    
6. Notes sur les Lepiotes. 2. Partie Generale. 1945. M. Locquin. Bulletin Mensuel de la Societe linneenne de Lyon 14: 85-89    
7. Notes on some Leucocoprinoid fungi from Britain. 1989. Derek A. Reid. Mycol. Res. 93 (4): 413-423    

物种描述

文献中物种描述

  • 以下描述摘自相关参考文献,由文字识别而来,请以原文为准。
  • 部分文献可能存在描述过时、描述错误或者描述片面等情况,仅供参考。
  • 外文描述的参考翻译系AI翻译或人工翻译,仅供参考。
4. Leucocoprinus ianthinus ( Cooke D Sacc .) Locq . in Bull . mens . Soc . linn . Lyon 14:94.1945.- Fig .53.
Agaricus ianthinus Cooke in Grevillea 16:101.1888, non Agaricus janthinus Fr .: Fr .,1821; Lepiota ianthina ( Cooke O ) Sacc ., Syll . Fung .9: 10.1891; Leucocoprinus ianthinus ( Cooke Ø Sacc .) Mohr in Boletus 18:48.1992( superfluous ).- Lepiota lilacinogranulosa P . Henn . in Verh . bot . ver . Prov . Brandenb .40:145.1898; Leucocoprinus lilacinogranulosus ( P . Henn .) Locq . in Bull . mens . Soc . linn . Lyon 12:94.1943; Leucocoprinus lilacinogranulosus ( P . Henn .) M . Bon et al . in Doc . mycol .19(75):53.1989( superfluous ); Hiatula cepaestipes var . lilacinogranulosa ( P . Henn .) Heim & Romagn . in Bull . trimest . Soc . mycol . Fr .50:184.1934.
SEL . ICON .- Heim & Romagn . in Bull . trimest . Soc . mycol . Fr .50: pl .11, fig .1.1934( as Hiatula cepaestipes var . lilacinogranulosa ); Migl . et al . in Riv . Micol .32:21.1989( as L . lilacinogranulosus );
SEL . DESCR .& FIGS .- Babos in Agarica 6(12):212-213. fig .14.1985( as L . lilacinogranulosus ): Candusso & Lanzoni , Lepiota :484-486, fig .107.1990( as L . lilacinogranulosus ); G . Fischer in Ulmer Pilzfl .1:136.1986; Heim & Romagn . in Bull . trimest . Soc . mycol . Fr .50:184-185, fig .12.1934 tas Hiatula cepaestipes var . lilacinogranu - losa ): Migl . et al . in Riv . Micol .32:20-23.1989( as L . lilacinogranu - losus ); D . Reid in Mycol . Res .93:418.1989( as L . ianthinus and L . lilacinogranulosus resp .).
VERN . NAME - Lilakorrelige plooiparasol .
Pileus in expanded specimens 17-62 mm , when young closed and paraboloid with flattened apex to ovoid , expanding to broadly cam - panulate , conical - campanulate and plano - convex with big , trun - cate umbo . and uplifted margin . at centre finely - squamulose felted , purplish , reddish brown ( K .& W .8E6,9D6,7E5), around cen -tre with small purplish fibrils , especially on the ridges , but mostly white or cream especially in outer half , and there without purplish fib - rils , costate - plicate up to 2/3 of radius or up to umbo , at utmost mar - gin yellowing with age , easily radially splitting , very thin - fleshed . Lamellae , L =35-55,1=(0-)1(-2), moderately distant to moderately crowded , free and remote from stipe , segmentiform - linear to ventri - cose , narrow , up to 3 mm wide , sometimes anastomosing - furcate , white with vague lilacinous sheen on some places when older , with finely flocculose white edge . Stipe 35-70(-150) x 2-5 mm , broaden - ing downwards and there bulbous - clavate , hollow , in apical part yel - lowish white , shiny , lengthwise fibrillose , below annulus white , or pale yellowish white with age , in lower half and at at bulb with purplish - violaceous fibrils to complete lilac covering ( e . g .11C4), with white tomentum at bulb itself , discolouring yellow when bruised . Annulus ascending , and slightly flaring , white , with purplish margin , evanes - cent . Context very thin and soft in pileus , white , concolorous with sur - face in stipe ( but not lilac - tinged ). Smell of cut basidiocarp fungoid . strong , slightly astringent . Taste fungoid . Spore print probably pale lilacinous to white .
Spores 8.0-12 x 5.5-7.5μm , on average 9.4-10.4 x 6.5-6.7μm . Q =1.25-1.7. Qav =1.4-1.55, in side - view ellipsoid - amygdaliform . some with apical papilla , in frontal view ellipsoid to slightly ovoid . ick - walled , with germ pore and hyaline cap over pore , dextrinoid , congophilous , cyanophilous , with pink inner wall and tract in Cresyl Blue . Basidia 17-42 x 8.0-11μ m .4- spored , rarely 2- spored , sur - rounded by 4-6 pseudoparaphyses . Lamella edge sterile ; cheilocystidia 27-80 x 9.0-35 μm . varying from cylindrical , utriform to broadly clavate , thin - walled , colourless . Pleurocystidia absent . Pileus covering made up of irregular chains of cylindrical to ellipsoid to globose ele - ments , with adnate to ascending elements , ranging from 15-35 x 5.0-9.0 μm to 25 um in diam .: pigment intracellular and lilac - brown to greyish lilac . Stipitipellis a cutis of cylindrical colourless hyphae ,4.0-13μm in diam ., with at base of stipe similar elements as on pileus sur - face . Clamp - connections absent .
HABITAT & DISTR .- In small groups , saprotrophic and terrestrial in pots of plants , in greenhouses . Rather rare in the Netherlands , found throughout the year , widespread in Europe , probably of tropical origin .
The name L . ianthinus is used for the species , more commonly known as L . lilacinogranulosus , following Bon ( Fl . mycol . Eur .3, L é piotes :112.1993); Reid ( in Mycol . Res .93:418.1989) gave arguments for and against this synonymy , but he did not reach a conclusion .
Reid ( in Mycol . Res .93:420.1989) distinguished L . lilacinogran - ulosus var . subglobisporus with smaller , subglobose spores ; this vari - ety has not yet been recorded from the Netherlands . Zuccherelli & Migliozzi ( in Boll . Gruppo micol . G . Bres ., n . S .,41:197-203.1998) reported on its occurrence in Italy .
Specimens with grey - lilac to lilac - brown tinges and narrowly lageniform to cylindrical elements on pileus and stipe are described under L . cepaestipes .

参考翻译:

成熟标本菌盖直径17–62毫米,幼时闭合,呈顶部扁平的拟球形至卵形,后展开为宽钟形、圆锥-钟形至平凸形,具大的截头状凸起,边缘上翘。中央区域具细小的鳞片状绒毛,呈淡紫色、红褐色(K. & W. 色卡 8E6, 9D6, 7E5),中央周围有淡紫色细纤毛,尤其在棱纹上,但大部分区域为白色或奶油色,尤其是在外部半部,此处无淡紫色纤毛。菌盖具肋状条纹,可达半径的2/3或直至凸起处。最边缘随年龄增长变黄,易沿径向开裂,菌肉极薄。菌褶数量 L = 35–55,短菌褶数量 l = (0–)1(–2),间距中等稀疏至中等密集,离生,远离菌柄,呈分段-线形至腹鼓状,窄,宽可达3毫米,有时分叉或呈网状,白色,老熟时局部隐约呈现淡紫色光泽,边缘具细絮状白边。菌柄 长35–70(–150)毫米,粗2–5毫米,向下渐粗,基部呈球棒状,中空。顶部近淡黄白色,有光泽,具纵向纤毛;菌环下方白色或随年龄增长呈浅黄白色;下半部及基部具淡紫色至紫罗兰色纤毛,甚至完全覆盖成淡紫色(11C4),基部本身有白色绒毛。受伤处变黄。菌环上位,向上略微展开,白色,边缘带淡紫色,易消失。菌盖菌肉处极薄且柔软,白色;菌柄处菌肉与表面颜色相同(但不带淡紫色调)。新鲜子实体切开时有明显的真菌气味,强烈,略带涩味。口味真菌味。孢子印可能为淡紫白色至白色。
孢子8.0–12 × 5.5–7.5 微米,平均 9.4–10.4 × 6.5–6.7 微米。Q值(长宽比) = 1.25–1.7,平均Q值 = 1.4–1.55。侧面观为椭圆-杏仁形,部分具顶端乳突;正面观为椭圆形至略卵圆形。壁厚,具发芽孔,孔上有透明盖帽。遇碘液呈糊精样反应,遇刚果红呈阳性,遇棉蓝呈蓝色。在甲苯胺蓝中内壁和纹路呈粉红色。担子17–42 × 8.0–11 微米,具4孢,罕见2孢,周围被4–6个周细胞包围。菌褶边缘不育。褶缘囊状体 27–80 × 9.0–35 微米,形态多变,从圆柱形、烧瓶形至宽棒状,薄壁,无色。侧生囊状体缺失。菌盖表皮由不规则链状的圆柱形至椭圆形至球形细胞构成,细胞贴生至直立,大小范围从 15–35 × 5.0–9.0 微米至直径达25微米。色素为细胞内,呈淡紫褐色至灰淡紫色。菌柄表皮为圆柱形无色菌丝组成的皮层,直径4.0–13微米。菌柄基部有与菌盖表面相似的结构。锁状联合缺失。
生境与分布:小群生长,腐生,陆生,常见于盆栽植物内及温室中。在荷兰较为罕见,全年可见,广泛分布于欧洲,可能起源于热带。
该物种学名 L. ianthinus 被用于指代这个通常更广为人知的 L. lilacinogranulosus 的物种,这一处理遵循 Bon (Fl . mycol . Eur .3, Lépiotes112. 1993) 的观点;Reid (Mycol. Res.93:418. 1989) 对是否合并正反两方面都给出了论据,但未得出结论。
Reid (Mycol. Res.93:418. 1989) 区分出 L. lilacinogranulosus var. subglobisporus,其孢子更小、近球形;此变种在荷兰尚无记录。Zuccherelli & Migliozzi (Boll . Gruppo micol . G. Bresadola, 新系列,41:197-203. 1998) 报道了其在意大利的分布。