指囊小蘑菇Micropsalliota digitatocystis

描述1 描述来源：A monograph of Micropsalliota in Northern Thailand based on morphological and molecular data
类型：原白（原始描述）
描述原文：
Micropsalliota digitatocystis R.L. Zhao, J.X. Li & M.Q. He, sp. nov.
Figure 2 Fungal Name: FN570747
Etymology: refers to the pleurocystidia with a digitate apical appendix.
Diagnosis: this new species is characterized by its brownish, fibrillose scales on pileus, stipe heavily covered white tomentose-flocculose, a needle-like pleurocystidia. Holotype: CHINA, Yunnan Province, Lushui County, Gaoligong National Natural Reserve, 8 July 2018, Zhi-Ling Ling, ZRL20180564 (HMAS290750).
Macroscopic description: Pileus 16–74 mm in diam., convex to applanate, subumbonate, margin crenate, surface dry, covered with fibrillose scales, slightly erect, denser at disc, scanty towards the margin, reddish brown, background white or light gray. Context up to 2 mm thick at disc. Lamellae free, crowded, with 4 series of lamellulae, 3–4 mm broad, grayish brown. Stipe 60–90 × 5–8mm, cylindrical with a slightly thickened base; surface above the ring white, smooth to fibrillose, decreasing with age, below the ring heavily covered with fibrils, white, yellowish-brown when bruised or cut. Annulus pendent, single, membranous, white, edge entire, persistent, up to 8–20 mm. Odor not distinctive. Microscopic description: Basidiospores 5.8–7.4 × 4–4.6 μm, [x = 6.9 ± 0.3 × 4.5 ± 0.2, Q = 1.5–1.8, Qm = 1.5 ± 0.085, n = 22], ellipsoid, sometimes amygdaliform, without germ pore, brown. Basidia 14.1–20.4 × 7.6–9.3 μm, clavate, hyaline, 4-spored. Cheilocystidia 37.4–52 × 9–16.4 μm, cylindrical to subclavate, slightly swollen at the middle, subcapitate to capitate, smooth, hyaline. Pleurocystidia present, with the similar size to basidia, nevertheless, carrying with a needle-like apex up to 4–5μm. Pileipellis composed of hyphae of 8–19 μm in diam., constricted at the septa, smooth, with internal brown pigment. Annulus composed of hyaline hyphae 5–18 μm in diam., smooth. KOH reaction: unknown.
Habit and habitat: solitary on soil in forest. Known distribution: China. Additional material examined: CHINA, Guangxi Zhuang Autonomous Region, Leye County, 7 August 2017, Mao-Qiang He, GX20170688 (HMAS290751).
Notes: Micropsalliota digitatocystis, M. globocystis, M. megarubescens, M. pseudoglobocystis and M. purpureobrunneola share similar morphological characteristics and they clustered together in the phylogenetic tree. However, M. digitatocystis is characterized by larger basidiomata (pileus diam. 18–74 mm, stipe 60–90 × 5–8mm), pileus margin dentate, floccose tomentose stipe bellow the annulus and digitate pleurocystidia. Micropsalliota globocystis and M. digitatocystis are morphologically very similar, but the former is characterized by its purple to purplish brown, grayish brown or reddish-brown pileus, and usually M. digitatocystis, presents light gray pileus. However, when M. digitatocystis has a reddish-brown pileus it also differs from M. globocystis in having digitate pleurocystidia. Micropsalliota megarubescens differs in having white to cream pileus becoming grayish brown with age and a strong red discoloration when touching or bruising. Micropsalliota pseudoglobocystis differs from M. digitatocystis, in having small basidiomata as well as small basidiospores (4.5–6 × 2.5–3.2μm) (Wei et al. 2015). Micropsalliota purpureobrunneola is distinguished from M. digitatocystis, in having small basidiomata (pileus 5–12 mm) and purple or purplish-brown scales (He et al. 2020).

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参考翻译：
针囊小蘑菇Micropsalliota digitatocystis R.L. Zhao, J.X. Li & M.Q. He，新种 Fungal Name: FN570747
词源：'digitatocystis'指的是其侧生囊状体顶部有指状的附属物。
该物种的主要特征是菌盖被褐色，细小纤维状的鳞片；菌柄被大量的绒毛或卷毛，由针状的侧生囊状体构成。主模式：中国云南泸水县，高黎贡自然保护区，2018年7月8日，采集人Zhi-Ling Ling，标本号ZRL20180564 (HMAS290750)。
宏观特征：菌盖宽16–74 mm，凸镜形至平展，中部近凸起，边缘锯齿状，表面干，底色白色或浅灰色，被大量细小纤维状鳞片，鳞片略翘起，朝周围渐变稀疏，红褐色。菌肉宽达2mm，菌盖中部更厚。菌褶离生，密集，小菌褶4轮，宽3-4mm，灰褐色。菌柄60–90 × 5–8mm，圆柱形，受伤变黄褐色，基部略膨大，菌环以上部分白色，光滑或被细纤维，随时间推移减少（鳞片减少？），菌环以下密被大量白色纤维质鳞片。菌环下垂，单层，膜质，白色，边缘完整，宿存，宽达8-20mm。气味不明显。显微特征：担孢子5.8–7.4 × 4–4.6 μm, [平均大小x = 6.9 ± 0.3 × 4.5 ± 0.2, 长宽比Q = 1.5–1.8, 平均长宽比Qm = 1.5 ± 0.085, n = 22]，椭圆形，有时杏仁形，无芽孔，褐色。担子14.1–20.4 × 7.6–9.3 μm，棒状，透明，4孢。褶缘囊状体37.4–52 × 9–16.4 μm,圆柱形至近棒状，中部略膨大，近头状或头状，光滑，透明。侧生囊状体存在，大小近似于担子，但顶部有4–5μm的针状附属物。盖皮由8–19 μm宽的菌丝组成，隔膜处变窄，光滑，有褐色胞内色素。菌环由宽5–18 μm的透明菌丝组成。
KOH反应：未知。
分布与生境：单生在土壤或林地表。已知分布在中国。额外研究标本：中国，广西壮族自治区，乐业县，2017年8月7日，采集人Mao-Qiang He，标本号GX20170688 (HMAS290751)。
讨论：针囊小蘑菇Micropsalliota digitatocystis, 球囊小蘑菇M. globocystis, 变红小蘑菇M. megarubescens, 假球囊小蘑菇M. pseudoglobocystis 和 M. purpureobrunneola几个物种形态特征较为相似，且在系统发育树上也聚在一起。针囊小蘑菇的特征是子实体较大，菌盖宽18–74 mm，菌柄60–90 × 5–8mm，菌盖边缘锯齿状，菌环以下的菌柄被大量鳞片，侧生囊状体指状。球囊小蘑菇的特征很相似，但其菌盖为紫色至紫褐色，灰褐色或红褐色，而针囊小蘑菇的菌盖底色一般为浅灰色，当其菌盖呈红褐色时也可以通过其指状的囊状体来区分。变红小蘑菇的菌盖白色至奶油色，老后变灰褐色，受伤后明显变红。假球囊小蘑菇的子实体更小，担孢子也更小(4.5–6 × 2.5–3.2μm) 。M. purpureobrunneola的子实体更小（菌盖5-12mm宽），被紫色或紫褐色的鳞片。