汉弗莱芝属Humphreya(网孢芝属)是灵芝家族中罕为人知的一个属,目前包含三个种(截至2024年2月4日),由Steyaert人在1972年建立,属名纪念灵芝属的解剖研究先驱者C. J. Humphrey。该属物种过去被置于广义假芝属Amauroderma s.lat.中。后经Steyaert对其微观特征的研究,发现A. lloydii担孢子具平截,其内孢壁具网状(蜂窝状)纹饰,故将A. longipes移入另成立的汉弗莱芝属Humphreya中。汉弗莱芝属成立之后,Decock和Welti等人也对汉弗莱芝属物种的孢子形态进行研究,基于C. coffeata和C. flavipora纵向或横向的脊状内孢壁纹饰与汉弗莱芝属其他物种网状或类网状内孢壁纹饰的差别,提出了关于C. coffeata和C. flavipora两物种在汉弗莱芝属中分类地位的质疑。后经Robledo等人结合系统发育学研究与孢子纹饰特征以及地理分布后,将原隶属于汉弗莱芝属的C. coffeata和C. flavipora移入新成立的脊孢芝属Cristataspora中。 a: Humphreya lloydii的担孢子; b: Humphreya endertii的担孢子。引自Species of Ganoderma and related genera mainly of the Bogor and Leiden herbaria 系统发育学方面,汉弗莱芝属物种现今仍未有可用的分子数据,其图片以及文献资料甚少,故目前汉弗莱芝属物种仅以其特殊的内孢壁网状纹饰来一定程度上证明其作为灵芝类真菌中一个独立分支的存在并与脊孢芝属物种区分开来(Sun Y.F. et al. 2022)
子实体一年生,具中生柄。菌盖平展至漏斗状;菌盖表面粘土色至乌黑色,不具漆状光泽,具辐射状分叉的沟纹,近中部具一深橄榄色的环纹。菌孔面白色,干后亦为白色;菌孔圆形;菌管与菌肉同色;菌肉浅黄色,干后可能为白色,近菌管层常具一条深橄榄色暗线;菌柄粘土色;菌柄表面不具漆状光泽。 分布:分布于西非地区。中国没有分布 生态:生于活立木(Paramacrolobium coeruleum)上 营养类型:寄生Parasitic 讨论:本种目前没有可用的分子数据。 相似物种:与部分灵芝属物种较为相似,但本种担孢子具网状内孢壁纹饰。 与Hu. eminii和Hu. endertii较为相似,但前者担孢子长度超过20μm,后者内孢壁纹饰为杂乱不规则的脊突。 原始描述:
Basidioma stipitate, mesopodial; pileus convex-infundibuliform, up to 120 mm in diam; dorsal surface with many radial, ramified folds, dullClay Color (Ridgway) to Sepia (Ridgway), in the latter case sometimes near the centre with narrow Dark Olive (Ridgway) rings. Pore surface white, usually remaining so upon drying; stipe may reach the formidable length of more than 500 mm, but is only 10-15 mm thick, usually about 200 mm long and 5-10 mm thick, dull Clay Color (Ridgway). Section: cutis very hard, dull dark olive, 60 μ thick; context varying in thickness with the folds of the surface, 1-4mm, probably white when fresh, Light Buff (Ridgway) when dry, often with a dark olive layer of melanoid substances near the tube layer of up to 1 mm thick; tubes up to 5-6 mm long, concolorous with context; under the cortex ofthe stipe sometimes a layer of melanoid substances which continue upwards in the pileus as a dark line over the tube layer. Cutis anamixodermiform, sharply defined, composed of the same hyphae as the context, the extremities somewhat inflated, impregnated by melanoid substances scarcely soluble in KOH. Context with periclinal or sometimes pantoclinal hyphae; hyphae 3-4 μ. thick, thick-walled, not producing skeletal hyphae. Pores round, 60-190-310 μ in diam.; dissepiments 10-95-280 μ thick; distance between axes about 285 μ. Spores ovoid, light yellowish, 12.5-16-20 X 8-10.5-15 μ, with crests in honeycomb pattern.